In late medieval Japan, Kai is a half-Japanese, half-English outcast who lives in the Ako domain, which is ruled by the benevolent Lord Asano Naganori.
When Kai was young, Asano found him lost in the forest and accepted him
into his household. Kai and Asano's daughter, Mika, eventually fall in
love despite Kai being scorned by her father's samurai due to his
lowborn status and mixed ancestry.
Before a planned visit from Shogun Tokugawa Tsunayoshi, Asano is visited by the Shogun's master of ceremonies, Lord Kira, who wants to take Ako for himself. Kira enlists the help of a shapeshifting kitsune
named Witch who sends a monster to kill Asano in the forest of Ako.
Asano's samurai struggle in their battle with the monster so Kai joins
in riding an abandoned horse. As the monster charges him, Kai recovers a
lost sword that he uses to slay it. Shortly thereafter he spots a white
fox with different coloured eyes
watching from the underbrush -- Witch in her fox form. Later during the
Shogun's visit, Kai sees that one of the Shogun's concubines has the
same two-colored eyes which leds him to assume that she was the fox from
the forest. He tries to warn Asano's principal counselor and samurai, Oishi, that she is a witch but Oishi doesn't believe him.
Later, Kira arranges a duel for the entertainment of the Shogun:
Kira's best warrior, the giant Lovecraftian Samurai, will battle a
warrior of Asano's choosing. However, before the duel begins, Witch uses
her magic to incapacitate Asano's combatant. Kai secretly dons his
armor and fights in his stead, but his disguise is revealed and the
Shogun orders him severely beaten as punishment. Later that night, Witch
uses her magic to make Asano believe that Kira is raping Mika, causing
him to attack an unarmed Kira in his delirium. Asano is sentenced to
death for attacking a Shogunate official (which is an executable offense), though the Shogun allows him to die with honor through seppuku.
The Shogun then gives Kira both the Ako domain and Mika, although he
grants Mika one year to mourn the death of her father before marrying
Kira. The Shogun also brands Oishi and his men ronin
and forbids them from seeking vengeance for Asano's death. To ensure
that the ronin don't interfere with his takeover plans, Kira imprisons
Oishi in an outdoor pit to break his spirit.
Nearly a year later, Oishi is released by Kira's men. He now knows
that Kira is guilty of treachery for using Witch's sorcery to cause
Asano's downfall. Oishi reunites with his family and asks his son
Chikara to aid him in reuniting the scattered ronin. They learn that Kai
has been sold into slavery and they rescue him from the fighting pits
of the Dutch colony of Dejima. Kai leads them to the Tengu
Forest, a mystical place he escaped from as a child, so that they can
find swords for themselves. Kai instructs Oishi to never draw his sword
while in the Tengu temple and continues alone to another room to face
the Tengu Master, who once trained Kai in their fighting ways. While Kai
confronts the Tengu Master in a battle of wills, Oishi watches an
illusion of his men being slaughtered by the Tengu, during which he
fights the urge to draw his own sword. With Kai and Oishi both
successful in their Tengu challenges, the ronin are given magical Tengu
swords.
Armed with their new weapons, the ronin plan to attack Kira on his
pilgrimage to a shrine where he seeks blessings for his wedding to Mika.
However, Kira's procession is a trap and the ronin are ambushed by
Kira's forces, led by Witch and the Lovecraftian Samurai. Several of the
ronin are killed, and Witch, thinking they are all dead, takes Oishi's
sword and presents it to Kira as a trophy. Witch later taunts Mika with
their deaths and attempts to manipulate her into committing suicide from
despair.
Oishi and Kai (having actually survived the attack) rally the
surviving ronin. Oishi and half of the ronin infiltrate Kira's (Asano's)
castle by disguising themselves as a band of traveling wedding
performers loyal to the memory of Lord Asano. With Kira's men distracted
during the performance, Kai and the remaining ronin scale the castle
walls, and the reunited group start to battle with Kira's men. While
Oishi fights Kira, Kai and Mika are attacked by Witch, who shape-shifts
into a dragon. Kai uses his magic sword and draws on the mystical powers
of the Tengu to finally kill her. Oishi then emerges with Kira's
severed head and Kira's men surrender.
After winning the battle, the ronin (including Kai) surrender
themselves to Shogunate authority and they are sentenced to death as
they explicitly violated the Shogun's prohibition on seeking revenge
against Kira. However, the Shogun finds that they followed the
principles of Bushido
in their actions and therefore restored their honor as samurai. Thus,
instead of being executed as criminals, the ronin are allowed to perform seppuku
so they can die with their honor restored. They are also given the
honor of burial with their master, Lord Asano. The Shogun gives Ako back
to Mika, and at the seppuku ceremony, he pardons Oishi's son Chikara so
that he may serve Ako and preserve such a noble samurai bloodline for
the country.
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